Peer Reviewed Publications
click pdf to access paper (*student or postbac technician, ^co-first authors)
2025
30. Gibbs, G.*, E. J. Resetarits, and S. L. Ziegler. 2025. Influence of salt marsh edge morphology on cross-boundary consumption
and prey density. Estuaries and Coasts 48(3): 77 doi: 10.1007/s12237-025-01513-4 pdf
29. Smith J.G., C. J. Lopazanski, C. M. Free, J. Brun, C. R. Anderson, M. H.Carr, J. Claudet, J.E. Dugan, J. G. Eurich , T. B. Francis,
D. A. Gill, S. L. Hamilton, K. Kaschner, D. Mouillot, P. T. Raimondi, R. M. Starr, S. L. Ziegler, D. Malone, M. L Marraffini, A. Parson-Field, B. Spiecker, M. Yeager, K. J. Nickols, and J. E. Caselle. 2025. Conservation benefits of marine protected areas accrue across a large ecologically connected network. Conservation Biology e14435 doi: 10.1111/cobi.14435 pdf
2024
28. Ziegler, S. L., W. R. James, M. Alber, and J.E. Byers. 2024. Seascape structure likely influences resource use of marsh-associated
estuarine consumers over a small spatial scale. Marine Ecology Progress Series 751: 1-12 doi: 10.3354/meps14755 pdf
27. Gross C. P., J. E. Duffy, K. A. Hovel, M. R. Kardish, P. L. Reynolds, C. Boström, K. A. Boyer, M. Cusson,J.Elköf, A. H. Engelen,
B. K. Eriksson, F. J. Fodrie, J. N. Griffin, C. M. Hereu, M. Hori, A. R. Hughes, M. V. Ivanov, P. Jorgensen, C. Kruschel, K. Lee, J. S. Lefcheck, P. Moksnes, M. Nakaoaka, M. I. O’Connor, J. L. Olsen, R. J. Orth, B. J. Peterson, H. Reiss, F. Rossi, J. L. Ruesink, E. E. Sotka, J. Thormar, F. Tomas, R. Unsworth, E. P. Voigt, M. A. Whalen, S. L. Ziegler, and J. J. Stachowicz. 2024. A latitudinal cline in the structure and taxonomic turnover of eelgrass epifaunal communities is associated with plant genetic diversity. Global Ecology and Biogeography 33:e13918 doi: 10.1111/geb.13918 pdf
26. Ziegler, S. L., M. J. Pierce, J. E. Byers. 2024. Invasive crab positively correlated with predatory crab species over a regional
scale. Biological Invasions 26: 3341-3351 doi: 10.1007/s10530-024-03390-7 pdf
25. Ziegler, S. L., W. E. Atencio, J. M. Carroll, J. E. Byers. 2024. High parasite prevalence in an ecosystem engineer correlated with
both local- and landscape-level factors. Oecologia 205: 423–435 doi: 10.1007/s00442-024-05581-4 pdf
24. Johnston, E. M. G. T. Waltz, R. Kosaka, E. M. Brauer, S. L Ziegler, E. T. Jarvis Mason, H. S. Glanz, L. Zaragoza, A. N. Kellum,
R. O. Brooks, B. X. Semmens, C. J. Honeyman, J. E. Caselle, L. F. Bellquist, S. L. Small, S. G. Morgan, T. J. Mulligan, C. L. Coscino, J. M. Staton, R. M. Starr, S.L. Hamilton, B. I. Ruttenberg and D. E. Wendt. 2024. Participation in Collaborative Fisheries Research Improves the Perceptions of Recreational Anglers towards Marine Protected Areas. Frontiers in Marine Science 11: 1330498 doi: 10.3389/fmars.2024.1330498 pdf
23. Ziegler S. L., R. O. Brooks, L. F. Bellquist, J. E. Caselle, S. G. Morgan, T. J. Mulligan, B. I. Ruttenberg, B. X. Semmens, R. M.
Starr, J. Tyburczy, D. E. Wendt, A. Buchheister, J. R. Marin Jarrin, C. Pasparakis, S. J. Jorgensen, J. A. Chiu, J. Colby, C. L. Coscino, L. Davis, F. de Castro, J. T. Elstner, C. Honeyman, E. T. Jarvis Mason, E. M. Johnston, S. L. Small, J. Staton, G. T. Waltz, B. Basnett, E. V. Satterthwaite, H. Killeen, C. D. Dibble, and S. L. Hamilton. 2024. Collaborative fisheries research reveals reserve size and age determine efficacy across a network of marine protected areas. Conservation Letters 17(2): e13000 doi: 10.1111/conl.13000 pdf
2023
22. Free C. M., J. G. Smith, J. Brun, T. B. Francis, J. G. Eurich, J. Claudet, J. E. Dugan, D. Gill, S. L. Hamilton, K. Kaschner, C. J.
Lopazanski, D. Mouillot, S. L. Ziegler, J. E. Caselle, and K. J. Nickols. 2023. If you build it, they will come: coastal amenities facilitate human engagement in marine protected areas. People and Nature 5(5): 1592-1609doi: 10.1002/pan3.10524 pdf
21. Yando E. S., S. F. Jones, W. R. James, D. D. Colombano, D. I. Montemayor, S. Nolte, J. L. Raw, S. L. Ziegler, L. Chen, D.
Daffonchio, M. Fusi, K. Rogers, and L. Sergienko. 2023. An integrative salt marsh conceptual framework for global comparisons. Limnology and Oceanography Letters 8(6): 830-849 doi: 10.1002/lol2.10346 pdf
20. Smith J.G., C. M. Free, C. J. Lopazanski, J. Brun, C. R. Anderson, M. H.Carr, J. Claudet, J.E. Dugan, J. G. Eurich , T. B. Francis,
S. L. Hamilton, K. Kaschner, D. Mouillot, P. T. Raimondi, R. M. Starr, S. L. Ziegler, K. J. Nickols, and J. E. Caselle. 2023. A marine protected area network does not confer community structure resilience to a marine heatwave across coastal ecosystems. Global Change Biology 29(19): 5634-5651 doi:10.1111/gcb.16862 pdf
19. Ziegler, S. L., J. M. Johnson*, R. O. Brooks, E. M. Johnston, J. L. Mohay, B. I. Ruttenberg, R. M. Starr,G. T. Waltz, D. E. Wendt,
and S. L. Hamilton. 2023. Marine protected areas, marine heatwaves, and the resiliency of nearshore fish communities. Scientific Reports 13:1405 doi:10.1038/s41598-023-28507-1 pdf
18. Clance, L. R.*^, S. L. Ziegler^, and F. J. Fodrie. 2023. Contaminants disrupt aquatic food webs via decreased consumer
efficiency. Science of the Total Environment 859(2): 160245 doi: 10.1016/j.scitotenv.2022.160245 ^co-first authors pdf
2022
17. Duffy J. E., J. J. Stachowicz, P. L. Reynolds, E. E. Sotka, K. A. Hovel, C. Boström, K.E. Boyer, M. Cusson, J. Eklöf, A. H. Engelen,
B. K. Eriksson, F. J. Fodrie, J. N. Griffin, C. M. Hereu, M. Hori, A. R. Hughes, M. V. Ivanov, M. Jahnke, P. Jorgensen, C. Kruschel, K. Lee, J. S. Lefcheck, P. Moksnes, M. Nakaoka, M. I. O'Connor, N. E. O'Connor, R. J. Orth, B. J. Peterson, H. Reiss, K. Reiss, J. P. Richardson, F. Rossi, J. L. Ruesink, S. T. Schultz, J. Thormar, F. Tomas, R. Unsworth, E. Voigt, M. A. Whalen, S. L. Ziegler, and J. L. Olsen. 2022. A Pleistocene legacy structures variation in modern seagrass ecosystems. Proceedings of the National Academy of Science 119 (32): e2121425119 doi: 10.1073/pnas.2121425119 pdf
16. Ziegler, S. L., R. O. Brooks, S. L. Hamilton, B. I. Ruttenberg, J. A. Chiu, R.T. Fields, G. T. Waltz, C. Shen, D. E. Wendt and R. M.
Starr. 2022. External fishing effort regulates the positive effects of no-take marine protected areas. Biological Conservation 269 (43):109546 doi: 10.1016/j.biocon.2022.109546 pdf
15. Gross C. P., J. E. Duffy, K. A. Hovel, M. R. Kardish, P. L. Reynolds, C. Boström, K. A. Boyer, M. Cusson, J. Elköf, A. H. Engelen,
B. K. Eriksson, F. J. Fodrie, J. N. Griffin, C. M. Hereu, M. Hori, A. R. Hughes, M. V. Ivanov, P. Jorgensen, C. Kruschel, K. Lee, J. S. Lefcheck, P. Moksnes, M. Nakaoaka, M. I. O’Connor, J. L. Olsen, R. J. Orth, B. J. Peterson, H. Reiss, F. Rossi, J. L. Ruesink, E. E. Sotka, J. Thormar, F. Tomas, R. Unsworth, E. P. Voigt, M. A. Whalen, S. L. Ziegler and J. J. Stachowicz. 2022. The biogeography of community assembly: latitude and predation pressure drive geographic variation in community trait distribution in a guild of marine epifaunal crustaceans. Proceedings of the Royal Society B doi:10.1098/rspb.2021.1762 pdf
2021
14. Ziegler, S. L., L. R. Clance*, A. R. McMains*, M. D. Miller*, and F. J. Fodrie. 2021. Influence of marsh island size on nekton
communities: intermediate optima rather than SLOSS. Marine Ecology Progress Series. doi:10.3354/meps13780 pdf
13. Hovel, K., J. E. Duffy, J. J. Stachowicz, P. Reynolds, C. Bostrom, K. Boyer, M. Cusson, F. J. Fodrie, K. Gagnon, C. Hereu, M. Hori,
P. Jorgensen, C. Kruschel, K. Lee, M. Nakaoka, N, O’Connor, F. Rossi, J. Ruesink, F. Tomas Nash and S. L. Ziegler. 2021. Joint effects of patch edges and habitat degradation on faunal predation risk in a widespread marine foundation species. Ecology. doi:10.1002/ecy.3316 pdf
12. Ziegler, S. L., R. Baker, S. C. Crosby, M. A. Barbeau, J. Cebrian, D. D. Colombano, R. M. Connolly, L. A. Deegan, B. L. Gilby,
D. Mallick, C. W. Martin, J. A. Nelson, J. F. Reinhardt, C. A. Simenstad, N. J. Waltham, T. A. Worthington, and L. P. Rozas. 2021. Geographic variation in salt marsh structure and function for nekton: finding commonality across multiple scales. Estuaries and Coasts. doi:10.1007/s12237-020-00894-y. pdf
11. Colombano, D.D., S. Y. Litvin, S. L. Ziegler, S. B. Alford, R. Baker, M. A. Barbeau, J. Cebrián, R. M. Connolly, C. A.Currin, L. A.
Deegan, J. S. Lesser, C. W. Martin, A. E. McDonald, C. McLuckie, B. H. Morrison, J. W. Pahl, L. M. Risse, J. A. M. Smith, L. W. Staver, R. E. Turner, and N.J. Waltham. 2021. Climate change implications for tidal marsh food webs and nekton support. Estuaries and Coasts. doi: 10.1007/s12237-020-00891-1 pdf
10. Waltham N. J., C. Alcott, M. A. Barbeau, J. Cebrian, R. M. Connolly, L. A. Deegan, K. Dodds, L. A. Goodridge Gaines, B. L.
Gilby, C. J. Henderson, C. M. McLuckie, T. J. Minello, G. S. Norris, J. Ollerhead, J. Pahl , J. F. Reinhardt, R. J. Rezek, C. A. Simenstad, J. A. M. Smith, E. L. Sparks , L. W. Staver , S. L. Ziegler and M. P. Weinstein. 2021. Tidal marsh restoration optimism in a changing climate and urbanizing seascape. Estuaries and Coasts. doi: 10.1007/s12237-020-00875-1 pdf
9. Ziegler, S. L., M. D. Miller*, C. S. Smith, and F.J. Fodrie. 2021. Abiotic cycles mediate the strength of cross-boundary
consumption within coastal food webs. Estuaries and Coasts 44: 1147-1156. doi:10.1007/s12237-020-00829-7 pdf
8. Gilby B. L., M. P. Weinstein, R. Baker, J. Cebrian, S. B. Alford, A. Chelsky, D. D. Colombano, R. M. Connolly, C. A. Currin, I. C.
Feller, A. Frank, J. A. Goeke, L. A. Goodridge Gaines, F. E. Hardcastle, C. J. Henderson, C. W. Martin, A. E. McDonald, B.H. Morrison, A.D. Olds, J. S. Rehage, N. J. Waltham, and S. L. Ziegler. 2020. Human actions alter tidal marsh seascapes and the provision of ecosystems services. Estuaries and Coasts. doi:10.1007/s12237-020-00830-0 pdf
2020
7. Baker R., M. D. Taylor, K. W. Able, M. W. Beck, J. Cebrian, D. D. Colombano, R. M. Connolly, C.
Currin, L. A. Deegan, I. C. Feller, B. L. Gilby, M. E. Kimball , T. J. Minello, L. P. Rozas, C. Simenstad, R. E. Turner, N. J. Waltham, M. P. Weinstein, S. L. Ziegler, P. S. E. zu Ermgassen, C. Alcott, S. B. Alford , M. Barbeau, S. C. Crosby, K. Dodds , A. Frank, J. Goeke, L. A. Goodridge Gaines, F. E. Hardcastle, C. J. Henderson, W. R. James, M. D. Kenworthy, J. Lesser, D. Mallick, C. W. Martin, A. E. McDonald, C. McLuckie, B. H. Morrison, J. A. Nelson, G. S. Norris, J. Ollerhead, J. W. Pahl, S. Ramsden, J. S. Rehage, J. F. Reinhardt, R. J. Rezek, L. M. Risse, J. A. M. Smith, E. L. Sparks, and L. W. Staver. 2020. Fisheries rely on threatened salt marshes. Science 370: 670-671. doi: 10.1126/science.abe9332 pdf
6. Whalen, M. A., R. D. B. Whippo, J. J. Stachowicz, P. H. York, E. Aiello, T. Alcoverro, A. H. Altieri, C.
Bertolini, L. Benedetti-Cehcci, M. Bresch, F. Bulleri, P. E. Carnell, S. Cimon, R. M. Connolly, M. Cusson, M. S. Diskin, E. D’Souza, A. A. V. Flores, F. J. Fodrie, A. W. E. Galloway, L. C. Gaskins, O. J. Graham, T. C. Hanley, C. J. Henderson, C. M. Hereu, M. Hessing-Lewis, K. A. Hovel, B. B. Hughes, A. R. Hughes, K. M. Hultgrin, H. Jänes, D. S. Janiak, L. N. Johnston, P. Jorgensen, B. P. Kelaher, C. Kruschel, B. S. Lanham, K. Lee, J. S. Lefcheck, E. Lozano-Álvarez, P. I. Macreadie, Z. L. Monteith, N. E. O’Connor, A. D. Olds, J. K. O’Leary, C. J. Patrick, O. Pino, A. G. P. Poore, M. A. Rasheed, W. W. Raymond, K. Reiss, O. K. Rhoades, M. T. Robinson, P. G. Ross, F. Rossi, T. A. Schlacher, J. Seemann, B. R. Silliman, D. L. Smee, M. Thiel, R. K. F. Unsworth, B. I. van Tussenbroek, A. Vergés, M. E. Yeager, B. K. Yednock, S. L. Ziegler, and J. E. Duffy. 2020. Climate drives the geography of marine consumption by changing predator communities. Proceedings of the National Academy of Science. doi:10.1073/pnas.2005255117 pdf
2015-2019
5. Ziegler, S. L., K. W. Able, and F. J. Fodrie. 2019. Dietary shifts across biogeographic scales alter spatial
subsidy dynamics. Ecosphere. 10 (12): e02980. doi:10.1002/ecs2.2980 pdf
4. Ziegler, S. L., J. H. Grabowski, C. J. Ballie, and F. J. Fodrie. 2018. Effects of landscape setting on
oyster reef structure and function largely persist more than a decade post-restoration. Restoration Ecology. 26 (5): 933-942. doi:10.1111/rec.12651 pdf
3. McCann, M. J., K. W. Able, R. R. Christian, F. J. Fodrie, O. P. Jensen, J. J. Johnson, P. C. López-Duarte,
C. W. Martin, J. A. Olin, M. J. Polito, B. J. Roberts, and S. L. Ziegler. 2017. Key taxa in food web responses to stressors: The Deepwater Horizon Oil Spill. Frontiers in Ecology and the Environment. 15(3): 242-149. doi:10.1002/fee.1474 pdf
2. Duffy J. E., S. L. Ziegler, J. Campbell, P. Bippus,* and J. S. Lefcheck. 2015. Squidpops: A simple tool
to crowdsource a global map of marine predation intensity. PLoS ONE 10(11): e0142994. doi:10.1371/journal.pone.0142994 pdf
1. Allen J. D., A. F. Armstrong and S. L. Ziegler. 2015. Environmental induction of polyembryony in
echinoid Echinoderms. Biological Bulletin 229: 221–231. doi:10.1086/bblv229n3p221 pdf
Publications in Review or Revision
1. Ziegler, S.L., J.S. Lesser, A.R. McMains*, P.C. Lopez-Duarte, J.A. Olin, M.J. Polito, and F.J. Fodrie. (In Review). Variation in cross
habitat energy flows to mobile estuarine consumers along a latitudinal gradient. Oikos
2. Pierce, M.J., T. Lynn, J. Grissett, S. L. Ziegler, S. Williams, S. Reddy, S. Dong, C. Bowser, M. Alber, S. C. Pennings, D. Mishra,
A. C. Spivak, and J. E. Byers. (In Revision). Scaling the spatial and temporal effects of wrack disturbance across a salt
marsh landscape. Marine Ecology Progress Series
Technical Reports
3. Caselle, J.E., K.J. Nickols, J.G. Smith, C. Lopanzanski, J. Bruen, C. Free, C. Anderson, M. Carr, J. Claudet, J. Dugan, J. Eurich,
T. Francis, D. Gill, S. Hamilton, K. Kaschner, D. Mouillot, P. Raimondi, R. Starr, and S.L. Ziegler. 2023. A Synthesis of ecological and social outcomes from the California MPA network. California Ocean Protection Council. doi:10.25494/P6430V
2. Hamilton, S.L., R. Starr, D. Wendt, B. Ruttenberg, J. Caselle, B. Semmens, L. Bellquist, S. Morgan, T. Mulligan, J. Tyburczy,
S. Ziegler, R. Brooks, G. Waltz, E. Mason, C. Honeyman, S. Small and J. Staton. 2022. California Collaborative Fisheries Research Program (CCFRP) – Monitoring and Evaluation of California Marine Protected Areas. CA Sea Grant. pdf
1. Starr, R. M., J. Caselle, A. Kahn, A. Lauermann, J. Lindholm, B. Tissot, S. Ziegler, C. Bretz, P. Carlson, K. Cieri, J. Hoeke,
C. Jainese, G. Martel, S. McDermott, J. Mohay and P. Salinas-Ruiz. 2022. Monitoring and Evaluation of Mid-Depth Rock Ecosystems in the California MLPA Marine Protected Area Network. CA Sea Grant. pdf